# 4D Molecular Therapeutics funding: rounds and investors > 4D Molecular Therapeutics has raised $307M across 3 rounds since 2015. Investors include Perceptive Advisors, Janus Henderson Investors and Casdin Capital. Source: https://millionrounds.com/company/4d-molecular-therapeutics (Millionrounds; every figure on the page links to its filing or article) United States · [Funded startup](https://millionrounds.com/companies) # Who invested in *4D Molecular Therapeutics* 3 rounds we have observed, with the 4 investors named in them. Every figure came from a named source, and where the company reported a raise to the SEC itself, that is shown beside it. Biotech Raised 5 months ago Funding summary Updated 26 Oct 2026 **$307M** raised across 3 rounds since 2015, from 3 sources **$7.0M** Venture **$100M** Round **$200M** Debt - **Latest round:** Debt, $200M Jun 2026 - **Step-up i:** 2× - **Investors:** 4 See who backed them ## The funding story Each circle is a round, placed by date and sized by amount. The line above is the running total. 2015 2017 2019 2021 2023 2025 **$7.0M** Venture **$100M** Round **$200M** Debt - **First round:** Venture, $7.0M - **Biggest round:** $200M - **Usual gap between rounds:** 10.3 years ## The rounds - **Jun 2026** Debt **$200M** *1 new* *3 returning* [Perceptive Advisors](https://millionrounds.com/fund/perceptive-advisors) [Janus Henderson Investors](https://millionrounds.com/fund/janus-henderson-investors) [Casdin Capital](https://millionrounds.com/fund/casdin-capital) [Hercules Capital](https://millionrounds.com/fund/hercules-capital) Source recorded - **Nov 2025** Round **$100M** *3 new* [Perceptive Advisors](https://millionrounds.com/fund/perceptive-advisors) [Janus Henderson Investors](https://millionrounds.com/fund/janus-henderson-investors) [Casdin Capital](https://millionrounds.com/fund/casdin-capital) Source recorded - **Aug 2015** Venture **$7.0M** No investor was named in the report. From a historical funding record SEC filing: $7.0M sold, first sale Aug 2015 ## Founders and team From 4D Molecular Therapeutics's own website and the SEC filings it signed: 1 founder, 9 on the team. **Theresa Janke** Chief of Staff, Co-Founder *Named on its 2018 Form D* - **Chris Simms** Chief Commercial & Business Officer - **Kim Maplestone** Chief Clinical Operations Officer - **Kristian Humer** Chief Financial Officer - **Glenn P. Sblendorio** Board Member - **Nancy Miller-Rich** Board Member - **David Kirn** Executive Officer, Director i - **David Schaffer** Executive Officer, Director i - **Leisa Johnson** Executive Officer i - **Peter Francis** Executive Officer i On the board, as filed i *Charles Theuer* *Hoyoung Huh* *Margi McLaughlin* *Tony Yao* ## Who was in which round The busiest backers across the latest rounds. A filled square is a round they joined; a dark one, a round they led. **Venture** 2015 **Round** 2025 **Debt** 2026 [Perceptive Advisors](https://millionrounds.com/fund/perceptive-advisors) [Janus Henderson Investors](https://millionrounds.com/fund/janus-henderson-investors) [Casdin Capital](https://millionrounds.com/fund/casdin-capital) [Hercules Capital](https://millionrounds.com/fund/hercules-capital) ## Everyone who backed them [Perceptive Advisors](https://millionrounds.com/fund/perceptive-advisors) Asset manager **2** rounds [Janus Henderson Investors](https://millionrounds.com/fund/janus-henderson-investors) Private equity **2** rounds [Casdin Capital](https://millionrounds.com/fund/casdin-capital) VC firm **2** rounds [Hercules Capital](https://millionrounds.com/fund/hercules-capital) Venture debt **1** round Missing a round, or an investor who was in one? [Tell us](https://millionrounds.com/corrections?slug=4d-molecular-therapeutics). ## Also claim to have backed them These 2 firms list 4D Molecular Therapeutics as a portfolio company on their own websites, but no report of a round names them. They are the firms' own claims, kept apart from the rounds above. [Amzak Health](https://millionrounds.com/fund/amzak-health) Growth equity Per its own site [Longevity Vision](https://millionrounds.com/fund/longevity-vision) Family office Per its own site ## Quick answers about 4D Molecular Therapeutics ### Who invested in 4D Molecular Therapeutics? 4D Molecular Therapeutics's investors include Perceptive Advisors, Janus Henderson Investors, Casdin Capital and Hercules Capital. ### How much has 4D Molecular Therapeutics raised? 4D Molecular Therapeutics has raised $307M across 3 rounds since 2015. ### When did 4D Molecular Therapeutics last raise? 4D Molecular Therapeutics's most recent round was a Debt of $200M in Jun 2026. ### What was 4D Molecular Therapeutics's biggest round? Its most recent round, $200M in Jun 2026, is also its largest on record. ### Who backed 4D Molecular Therapeutics first? 4D Molecular Therapeutics's first round on record, in Nov 2025, was backed by Perceptive Advisors, Janus Henderson Investors and Casdin Capital. ### What stage is 4D Molecular Therapeutics at? 4D Molecular Therapeutics's latest named round is a Debt, in Jun 2026. ### Where is 4D Molecular Therapeutics based? 4D Molecular Therapeutics is based in the United States. ## Companies like 4D Molecular Therapeutics In the same sector, and raised in the last eighteen months. 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